Jonathan Roughgarden

Joan Roughgarden is an American theoretical ecologist and evolutionary biologist. She is professor of biology and of geophysics emerita at Stanford University and an adjunct researcher at the Hawaiʻi Institute of Marine Biology. She combined mathematical theory with field studies of Caribbean Anolis lizards and of larval recruitment in rocky-intertidal barnacles, wrote textbooks in ecological and evolutionary theory, and founded Stanford's Earth Systems Program.

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